1997-PhD thesis defence, University of the Basque Country, Spain: The role of inflammatory meditors in the regulation of receptor-mediated endocytosis by the hepatic sinusoidal endothelium.
1997-1999- Postdoctoral fellowship. YALE University (USA). Project: Regulation of lysosomal exocytosis in mammalian cells.
2000-2002- Research associate: University of Navarra (Spain). Project: Expression of Anion exchanger 2 (AE2) in hepatic epithelial cells and its role in primary biliary cirrhosis.
20002-2005: Dominion-Pharmakine Ltd. (Bilbao, Spain). Director of the Cell Biology Laboratory.
2005-2011: Associate Professor, University of Tromsø, Department of Clinical Medicine. Research Group: Bone and Joint Research Group.
2007-2009: Orthogenics TM, Research Park, Tromsø. Chair of the R&D group.
2012: Professor in Cell Biology, University of Tromsø, Deparment of Clinical medicine.